Ornithischia Classification
Order Ornithischia
Traditional classification
The following is based mainly on Carroll (1988), as the most recent comprehensive linnaean classification of the Vertebrates, with more recent groups added. Each taxon is listed by rank, name, and original author, and followed by a short description. Some additions have also been made from Paul 2010 and Holtz 2011. Linnaean names may not match their cladistic equivalents. New taxa are indicated by inverted commas. MAK120308 120312
Infraclass
Archosauromorpha Huene, 1946 (
cont.)
Order
Ornithischia Seeley, 1888
"bird-hipped" dinosaurs
Suborder
Ornithopoda Marsh, 1881
generalised bipedal, herbivorous dinosaurs, mostly a "wastebasket taxon" for anything taht doesn't fit in the other groups. L Trias to L Cret, Cosm.
Family Pisanosauridae Casamiquela, 1967
the most primitive group of ornithischians, monospecies only, L Tr of S Am
Family
Fabrosauridae Galton, 1972
small bipedal generalised ancestral ornithischians, E Jur to M Jur of S Afr, EAs.[1]
Family
Heterodontosauridae Romer, 1966
small bipedal early specialised forms, E Jur to E Cret of S Afr, Eur, EAs.
Family
Hypsilophodontidae Dollo, 1882
small bipedal, cursorial, generalised ornithopods, ancestral to later groups, M Jur to L Cret. Cosm.[2]
Families not specified -
a number of iconic types such as Muttaburrasaurus ( M Cret of Aus) and Tenontosaurus (M Cret of N Am) that don't fit easily into the established groups, perhaps require monospecific families or subfamilies
Family Rhabdodontidae
medium-sized ornithopods that represent a distinct evolutionary line - L Cret of Eur
Family
Dryosauridae small to medium sized, bipedal, cursorial, ornithopods, intermediate between hypsilophodonts and iguanodonts, Monogeneric. L Jur of E Afr, N Am.
Family
Iguanodontidae Cope, 1869
large to very large, bipedal and faculatively quadrapedal, ornithopods, intermediate between dryosaurs and hadrosaurs. L Jur to L Cret, Eur, N Am, As.
Family Hadrosauridae
"duck-billed" dinosaurs, large to very large, bipedal and faculatively quadrapedal, ornithopods, highly specialsed dental batteries, frequently with head crests. E to L Cret, N Am, As, Eur, SAm.
Subfamilies not specified -
a large assortment of various primitive and unspecialised Hadrosaurus, E to L Cret, N Am, As, Eur, SAm.
Subfamily
Saurolophinae -
flat-headed, broad-Snouted hadrosaurs - L Cret of N Am & E As
Tribe Maiasaurini—
unspecialised, primitive, broad-snouted duckbilled dinosaurs - L Cret of N Am
Tribe Saurolophini—
spike-crested duckbilled dinosaurs - L Cret of N Am & E As
Tribe Edmontosaurini—
very broad-snouted (classic "duck bill" dinosaurs - L Cret of N Am & E As
Subfamily
Lambeosaurinae -
crested hadrosaurs - L Cret of N Am
Tribe Saurolophini -
tube crested lambeosaurines
Tribe Corythosaurini (=Lambeosaurini) -
helmet crested lambeosaurines
Suborder unspecified
[3]
Family
Scelidosauridae Cope, 1869 -
Ancestral armoured types - mostly Early Jurassic. Quadrupedal, ancestral armoured dinosaurs. Includes light running forms and large heavily armed types. Length 1 to 4.5 meters E Jur of Eur, NAm, As
Suborder
Stegosauria Marsh, 1877 -
Plated dinosaurs - spines along the back and tail for protection, plates served a thermoregulatory purpose. Quadrupedal, short forelimbs, slow-moving. Length 3 to 10 meters M Jur to E Cret of N Am, Eur, As, Afr
Family Huayangosauridae -
Primitive plated dinosaurs; M Jur of E As
Family Stegosauridae Marsh, 1877
advanced plated dinosaurs; M Jur to E Cret of N Am, Eur, As, Afr
Subfamily Kentrosaurinae -
small plates, shoulder spines, spikes along back and tail - usually given family rank (Kentrosauridae); may be monotypic - L Jur of Afr, ?Eur
Subfamily Dacentrurinae -
tail spikes like Stegosaurus, otherwise resembles Kentrosaurus L Jura of Eur
Subfamily Stegosaurinae -
advanced stegosaurs, alternatinbg paired plates, no shoulder spines; L Jur to E Cret of NAm & EAs
Suborder
Ankylosauria Osborn, 1923
heavily armoured dinosaurs, 2 to 10 meters long, quadrupedal, slow moving, and fed on soft vegetation.
Family Polocanthidae -
ancestral ankylosaurs - L Jur to L Cret of Eur, N Am, EAs & Ant.
Family
Nodosauridae Marsh, 1890 -
with spikes on the shoulders - E to L Cret of Eur & N Am
Family
Ankylosauridae Brown, 1908 -
with a war club at the end of the tail - E to L Cret of E As & N Am
Suborder
Pachycephalosauria Sternberg, 1945
(Only one family; Homalocephalidae Dong, 1974 is comprised of juvenile pachycephalosaurids only)
Family
Pachycephalosauridae Sternberg, 1945
dome headed dinosaurs, superficially resemble the ornithopods; bipedal, small to medium-sized, with a bony "battering ram" head. L Cret of NAm & E As
Suborder
Ceratopsia Marsh, 1890 horned dinosaurs, small to large parrot-beaked dinosaurs,
Family Chaoyangsauridae Zhao, Cheng, & Xu, 1999
ancestral horned dinosaurs (or ancestral marginocephalia?) L Jur of EAs
Family
Psittacosauridae Osborn, 1923
parrot-beaked dinosaurs, stocky animals about about 1.5 meters long, hornless, bipedal and faculatively quadrapedal, E Cret of EAs
Family
Protoceratopsidae Granger & Gregory, 1923
small to medium sized ancestral hornless forms, bipedal or quadrapedal L Cret of EAs & NAm
Subfamily Leptoceratopsinae -
short frilled protoceratopsids - L Cret of E AS & N Am
Subfamily Bagaceratopinae -
Small-horned, lump-nosed Frilled Dinosaurs - L Cret of Eur & E As
Subfamily Protoceratopsinae -
advanced, deep-tailed protoceraptopsids - L Cret of E As
Family Ceratopsidae Marsh, 1890
large to gigantic horned dinosaurs, quadrapedal, rhino-like L Cret of NAm
Subfamily
Centrosaurinae -
short-frilled horned dinosaurs
Subfamily
Chasmosaurinae -
long-frilled horned dinosaurs
Hybrid classification
This section is part of a discontinued and impractical attempt to integrate the rank-based Linnaean and the phylogeny-based Cladistic systems . We have distinguished between monophyletic and paraphyletic taxa (or clades and grades), as indicated by the following key regarding prefixes:
For the phylogenetic arrangement, see the dendrogram page MAK120308 120312
Parainfraclass
Archosauromorpha Huene, 1946 (
cont.)
Holoorder
Ornithischia Seeley, 1888
"bird-hipped" dinosaurs
Holosuborder
Ornithopoda Marsh, 1881
generalised bipedal, herbivorous dinosaurs, mostly a "wastebasket taxon" for anything taht doesn't fit in the other groups. L Trias to L Cret, Cosm.
Monofamily Pisanosauridae Casamiquela, 1967
the most primitive group of ornithischians, monospecies only, L Tr of S Am
Parafamily
Fabrosauridae Galton, 1972
small bipedal generalised ancestral ornithischians, E Jur to M Jur of S Afr, EAs.
Holofamily
Heterodontosauridae Romer, 1966
small bipedal early specialised forms, E Jur to E Cret of S Afr, Eur, EAs.
Parafamily
Hypsilophodontidae Dollo, 1882
small bipedal, cursorial, generalised ornithopods, ancestral to later groups, M Jur to L Cret. Cosm.
Families not specified -
a number of iconic types such as Muttaburrasaurus ( M Cret of Aus) and Tenontosaurus (M Cret of N Am) that don't fit easily into the established groups, perhaps require monospecific families or subfamilies
Holofamily Rhabdodontidae
medium-sized ornithopods that represent a distinct evolutionary line - L Cret of Eur
Evofamily
Dryosauridae small to medium sized, bipedal, cursorial, ornithopods, intermediate between hypsilophodonts and iguanodonts, Monogeneric. L Jur of E Afr, N Am.
Parafamily
Iguanodontidae Cope, 1869
large to very large, bipedal and faculatively quadrapedal, ornithopods, intermediate between dryosaurs and hadrosaurs. L Jur to L Cret, Eur, N Am, As.
Holofamily Hadrosauridae
"duck-billed" dinosaurs, large to very large, bipedal and faculatively quadrapedal, ornithopods, highly specialsed dental batteries, frequently with head crests. E to L Cret, N Am, As, Eur, SAm.
Subfamilies not specified -
a large assortment of various primitive and unspecialised Hadrosaurus, E to L Cret, N Am, As, Eur, SAm.
Holosubfamily
Saurolophinae -
flat-headed, broad-Snouted hadrosaurs - L Cret of N Am & E As
Ambitribe Maiasaurini—
unspecialised, primitive, broad-snouted duckbilled dinosaurs - L Cret of N Am
Ambitribe Saurolophini—
spike-crested duckbilled dinosaurs - L Cret of N Am & E As
Ambitribe Edmontosaurini—
very broad-snouted (classic "duck bill" dinosaurs - L Cret of N Am & E As
Holosubfamily
Lambeosaurinae -
crested hadrosaurs - L Cret of N Am
Ambitribe Parasaurolophini -
tube crested lambeosaurines
Ambitribe Corythosaurini (=Lambeosaurini) -
helmet crested lambeosaurines
Suborder unspecified
Evofamily
Scelidosauridae Cope, 1869 -
Ancestral armoured types - mostly Early Jurassic. Quadrupedal, ancestral armoured dinosaurs. Includes light running forms and large heavily armed types. Length 1 to 4.5 meters E Jur of Eur, NAm, As
Suborder
Stegosauria Marsh, 1877 -
Plated dinosaurs - spines along the back and tail for protection, plates served a thermoregulatory purpose. Quadrupedal, short forelimbs, slow-moving. Length 3 to 10 meters M Jur to E Cret of N Am, Eur, As, Afr
Parafamily Huayangosauridae -
Primitive plated dinosaurs; M Jur of E As
Holofamily Stegosauridae Marsh, 1877
advanced plated dinosaurs; M Jur to E Cret of N Am, Eur, As, Afr
Monosubfamily Kentrosaurinae -
small plates, shoulder spines, spikes along back and tail - usually given family rank (Kentrosauridae); may be monotypic - L Jur of Afr, ?Eur
Evosubfamily Dacentrurinae -
tail spikes like Stegosaurus, otherwise resembles Kentrosaurus L Jura of Eur
Holosubfamily Stegosaurines -
advanced stegosaurs, alternatinbg paired plates, no shoulder spines; L Jur to E Cret of NAm & EAs
Suborder
Ankylosauria Osborn, 1923 -
heavily armoured dinosaurs, 2 to 10 meters long, quadrupedal, slow moving, and fed on soft vegetation.
Evofamily Polocanthidae -
ancestral ankylosaurs - L Jur to L Cret of Eur, N Am, EAs & Ant.
Holofamily
Nodosauridae Marsh, 1890 -
with spikes on the shoulders - E to L Cret of Eur & N Am
Holofamily
Ankylosauridae Brown, 1908 -
with a war club at the end of the tail - E to L Cret of E As & N Am
Suborder
Pachycephalosauria Sternberg, 1945
(Only one family; Homalocephalidae Dong, 1974 is comprised of juvenile pachycephalosaurids only)
Holofamily
Pachycephalosauridae Sternberg, 1945
dome headed dinosaurs, superficially resemble the ornithopods; bipedal, small to medium-sized, with a bony "battering ram" head. L Cret of NAm & E As
Suborder
Ceratopsia Marsh, 1890 horned dinosaurs, small to large parrot-beaked dinosaurs,
Evofamily Chaoyangsauridae Zhao, Cheng, & Xu, 1999
ancestral horned dinosaurs (or ancestral marginocephalia?) L Jur of EAs
Evofamily
Psittacosauridae Osborn, 1923
parrot-beaked dinosaurs, stocky animals about about 1.5 meters long, hornless, bipedal and faculatively quadrapedal, E Cret of EAs
Evofamily
Protoceratopsidae Granger & Gregory, 1923
small to medium sized ancestral hornless forms, bipedal or quadrapedal L Cret of EAs & NAm
Evosubfamily Leptoceratopsinae -
short frilled protoceratopsids - L Cret of E AS & N Am
Evosubfamily Bagaceratopinae -
Small-horned, lump-nosed Frilled Dinosaurs - L Cret of Eur & E As
Evosubfamily Protoceratopsinae -
advanced, deep-tailed protoceraptopsids - L Cret of E As
Holofamily Ceratopsidae Marsh, 1890
large to gigantic horned dinosaurs, quadrapedal, rhino-like L Cret of NAm
Holosubfamily
Centrosaurinae -
short-frilled horned dinosaurs
Holosubfamily
Chasmosaurinae -
long-frilled horned dinosaurs
Notes
[1] either a wastebasket taxon for any really primitive ornithischians (apart from Pisanosaurus) or limited to one or two Early Jurassic species from southern Africa. if the former, the boundaries between the the Fabrosauridae and other stem groups such as the Scelidosauridae and the Hypsilophodontidae are very poorly defined
[2] another ancestral/generalised/wastebasket taxon; generally, all ornithopods beneath the level of dryosaurs. Like other paraphyletic (ancestral) taxa, hypsilophodonts are defined in terms of shared primitive features (plesiomorphies)
[3] Essentially, basal Thyreophora. Although monophyletic Thyreophora and Cerapoda are sometimes given subordinal rank, we have followed Carroll (1988) in retaining the five classic ornithischian suborders
Links: The major dinosaur families at Enchanted Learning; Taxon tree; Linnean Ornithischia, dinosaur mailing list
page MAK120308, last modified 120506