Subclass Anapsida
Order Chelonii Brogniart 1800 (Testudinata Oppel 1811, Testudines Batsch 1788, Chelonia of Romer etc)
Suborder "Odontochelydia"
Family †Odontochelyidae Li et al., 2008 (monotypal - †
Odontochelys)
Suborder †Proganochelydia, Romer, 1966
Family †Proganochelyidae Baur 1888 (
Proganochelys,
Chinlechelys Joyce et al. 2008)
Family †Australochelyidae Gaffney & Kitching, 1994 (
Australochelys and
Palaeochersis - paraphyletic)
Family †Proterochersidae Nopcsa, 1928 (monotypal -
Proterochersis, convergent with Pleurodira)
Suborder Cryptodira Cope, 1868 (= Cryptodiramorpha Lee 1995)
Infraorder unspecified
Family †Kayentachelyidae (Kayentachelys Gaffney et al. 1987 monotypic family)
Family †Indochelyidae Datta et al. 2000 (
Indochelys - monotypic family)
Family †Solemydidae (may be Ceratocryptodira)
Family †Kallokibotiidae Nopcsa 1923 (may be Ceratocryptodira)
Family unspecified
†
Condorchelys Sterli, 2008,
Eileanchelys
Infraorder †Ceratocryptodira Bour & Dubois 1986
Superfamily †Meiolaniodea Gaffney
Family †Meiolaniidae Lydekker, 1887 (horned turtles)
Family †Mongolochelyidae Sukhanov and Pozdnjakov (Mongolochelys - monotypal)
Family †Chelycarapookidae Warren 1969 (Chelycarapookus - monotypal)
Family unspecified
†
Chubutemys Gaffney et al. 2007
Family unspecified
Otwayemys Gaffney et al. 1998
Infraorder †Paracryptodira Gaffney, 1975 (= Amphichelydia Lydekker 1889) (semi-aquatic, terrapin-like)
Superfamily †Pleurosternoidea Hay, 1930 (= Basicryptodira Bour & Dubois 1986)
Family †Pleurosternidae Cope, 1868
Family †Glyptopsidae Marsh, 1890 (may or may not be synonymous with Pleurosternidae)
Superfamily †Baenoidea Williams, 1950
Incertae sedis
†
Arundelemys
†
Compsemys
Family †Baenidae Cope 1882
Subfamily †Neurankylidae Lambe, 1902 - here used for paraphyletic
Neurankylus lineage of
Lyson & Joyce 2009 includes monotypal †Trinithchelyinae Gaffeny, 1972
Subfamily: †Hayemydinae Gaffney, 1972 (monotypal -
Hayemys)
Subfamily †Palatobaeninae Gaffney, 1972 - here used for
Plesiobaena-
Palatobaena lineage of
Lyson & Joyce 2009
Subfamily †Eubaeninae Gaffney, 1972 - here used for
Eubaena-
Stygiochelys lineage of
Lyson & Joyce 2009
Infraorder Eucryptodira Gaffney 1975 (incl. Pancryptodira Joyce et al. 2004)
Parvorder unspecified
Superfamily †Plesiochelyoidea (paraphyletic, stem eucryptodires, all freshwater aquatic or marginal marine)
Family †Eurysternidae Dollo, 1886
Family †Thalassemydidae Zittel 1889 (monotypal -
Thalassemys Rütimeyer, 1859)
Family †Plesiochelyidae Rütimeyer, 1873 (Plesiochelys Rütimeyer, 1873, Hylaeochelys Lydekker, 1889, Portlandemys Gaffney 1975)
Family †Protostegidae Cope, 1872 (small to gigantic Cretaceous marine turtles, cosmop., convergent with Dermochelyidae)
Incertae sedis
†
Santanachelys Hirayama, 1998
†
Notochelone Owen, 1882
etc.
Subfamily †Desmatochelyinae Williston, 1894 (monotypal -
Desmatochelys - Late Cretaceous of N Am)
Subfamily †Chelospharginae Zanger 1953 (paraphyletic grade of small primitive types - Late Cretaceous of N Am, if includes
Rhinochelys Seeley 1869 then also European distribution)
Subfamily †Protosteginae Zangerl, 1953 (large to giant advanced forms - Late Cretaceous of N Am)
Clade †Angolachelonia Mateus et al. 2009 (= family "Angolocheloniidae"?)
Superfamily unspecified - (eucryptodires of Xinjiangchelyid grade, generally semi-aquatic, terrapin-like)
Family †Xinjiangchelyidae (paraphyletic ancestral forms, Mid Jurassic to Early Cretaceous of Asia)
Incertae sedis
Chengyuchelys
Brodiechelys
Larachelus
etc.
Parvaorder Centrocryptodira Gaffney and Meylan 1988 ((including Polycryptodira Gaffney and Meylan 1988)
Superfamily unspecified, paraphyletic or polyphyletic (testudines of Macrobaenid-Sinemydid grade, semi-aquatic, terrapin-like)
Family †Sinemydidae Yeh 1963 (primitive forms - Late Jurassic to Paleogene)
Family †Macrobaenidae Sukhanov 1964 (
Judithemys &
Macrobaena - advanced stem or crown centrocryptodires)
Incertae sedis
†
Hangaiemys
†
Hoyasemys Perez-Garcia et al. 2012
†
Wuguia Matzke et al. 2004
Superfamily Trionychoidea Fitzinger, 1826 (sensu Trionychoidae = Pantrionychia Joyce et al. 2004)
Epifamily †Adocoidae Cope, 1870 - shell sculpture in the form of pits, separated by ridges. Late Jurassic - Oligocene of Asia & North America.
Family †Adocidae Cope, 1870
Subfamily †Adocinae Cope, 1870
Subfamily †Shachemydinae Khosatzky in Nessov & Khosatzk y , 1977 -
Ferganemys &
Shachemys - Cretaceous of Asia
Family †Nanhsiungchelyidae Yeh, 1966 - shell sculpture in the form of large pits with high crests - Cretaceous of Asia and North America.
Epifamily Trionychoidae (softshell turtles and their ancestors Late Hurasisc to Recent)
Family Trionychidae Fitzinger, 1826 (softshell turtles)
Family Carettochelyidae Boulenger, 1887 (pignose turtles)
Superfamily Chelonioidea Baur 1893 (= Panchelonioidea Joyce et al. 2004) (sea turtles - Cretaceous to recent)
Family unspecified
†
Chitracephalus Dollo 1884 (small, semi-aquatic)
Incertae sedis
Subfamily †Allopleuroninae (Large sea turtles, Late Cretaceous of Europe, convergent with Protostegids and Dermatemydids)
Family †Toxochelyidae Baur 1895
Subfamily †Toxochelyinae (medium to large near shore marine types - Late Cretaceous of N Am)
Subfamily †Lophochelyinae (Late Cretaceous - may be Toxochelyids or Cheloniids)
Family Cheloniidae Oppel, 1811 (sea turtles)
Incertae sedis
†
Mexichelys
†
Euclastes
†
Gigantatypus - late Cretaceous
Subfamily †Eocheloniinae (ancestral types, Paleocene to Miocene)
Subfamily Cheloniinae (recent and related sea turtles - Miocene to Recent, Cosmop.)
Tribe Chelonini (Green sea turtles and their relatives; incl. Syllominae)
Tribe Carettini (Loggerheads and their relatives)
Family Dermochelyidae Fitzinger, 1843 (leatherback sea turtles - Late Cretaceous to Recent)
Superfamily Chelydroidea Baur 1893
Family Chelydridae Gray, 1831 (snapping turtles)
Family Platysternidae Gray, 1869 (big-headed turtle) (= Platysternoidea Baur 1893 )
Superfamily Kinosternoidea Holroyd and Hutchison 2002
Family Dermatemydidae Gray, 1870 (river turtles)
Family Kinosternidae Agassiz, 1857 (mud turtles)
Subfamily
Subfamily
Superfamily Testudinoidea Gray 1825
Family †Haichemydidae Sukhanov and Narmandakh 2006
Family †Lindholmemydidae Chkhikvadze, 1970
Family †Sinochelyidae Chkhikvadze, 1970 (incl. Peishanemydidae Nesov)
Family Emydidae Rafinesque, 1815 (pond, box, and water turtles)
Family Geoemydidae Theobald, 1868 ((Asian river turtles, Asian leaf turtles, Asian box turtles, and roofed turtles)
Family Testudinidae Batsch, 1788 (true tortoises - Late Paleocene to Recent)
Incertae sedis
†
Achilemys Hay, 1908
†
Hadrianus Cope, 1872
etc.
Subfamily Xerobatinae Gaffney & Maylan, 1988 (desert and burrowing tortoises, Late Eocene to Recent)
Subfamily Testudinidae (most tortoises, Paleogene to Recent)
Tribe Testudinini - typical tortoises
Tribe Megalochelyini - average to giant tortoises
Suborder Pleurodira Cope, 1864 (3 families) (incl. Eupleurodira Gaffney and Meylan 1988)
Superfamily unspecified
Family †Platychelyidae Bräm, 1965 (= Platychelira Gaffney et al. 2006)
Superfamily unspecified
Family †Dortokidae de Lapparent de Broin and Murelaga 1999
Superfamily unspecified
Family †Propleuridae Cope 1872
Superfamily unspecified
Family †Eusarkiidae Bergounioux 1952
Superfamily unspecified
Family Notoemydidae de Broin and de la Fuente 1993
Superfamily unspecified
Family †Araripemydidae
Superfamily unspecified
Family Chelidae Gray, 1825 (Austro-American sideneck turtles)
Superfamily Pelomedusoidea (incl Pelomedusoides (Broin 1988) )
Family †Bothremydidae (incl Bothremydinura Gaffney et al. 2006)
Family Pelomedusidae Cope, 1868 (African sideneck turtles)
Family Podocnemididae Cope, 1868 (Madagascan big-headed and American sideneck river turtles)