Palaeos Palaeos Chelonii
Vertebrates References

Chelonii: References

Abbreviated Dendrogram
ANAPSIDA ?  
|
o CHELONII
|--Odontochelys   
`--+--Proganochelys
   `--+--Proterochersis 
      `--+--Pleurodira
         |--Paracryptodira 
         `--o Eucryptodira
            |--Protostegidae
            `--+--Angolachelonia
               `--+--Xinjiangchelyidae
                  `--+==Sinemydidae
                     |--Trionychoidea
                     `--+--+--Chelonioidea
                         |  |--Chelydridae
                         |  `--Kinosternoidea
                         `--Testudinoidea
Contents

Overview
Chelonian origins
Testudines
Casichelydia
Derived Stem Testudines
Meiolanoidea
Pleurodira
Paracryptodira
Eucryptodira
Protostegidae
Centrocryptodira
Trionychoidea
Chelonioidea
Chelydridae
Kinosternoidea
Testudinoidea
Testudinidae
Dendrogram
References




References



Abel, Othenio. 1919. Die Stämme der Wirbeltlere. Verinigung Wissenschaftlicher Verleger. Berlin und Leipzig.
Pleurosternon

John Acorn, 2007. Deep Alberta: Fossil Facts and Dinosaur Digs. Edmonton: University of Alberta Press
Basilemys

Allendorf, F. W., R. F. Leary, P. Spruell & J. K. Wenburg. 2001. The problems with hybrids: setting conservation guidelines. Trends in Ecology and Evolution 16 (11): 613-622.
Tortoise Resurrection

Anon. January 2009, St. Catherines Island Sea Turtle Conservation Program. web page

Anquetin, J. (2009) A new stem turtle from the Middle Jurassic of the Isle of Skye, Scotland, and a reassessment of basal turtle relationships. Doctoral thesis, UCL (University College London). pdf, mirror.
Chelonian phylogenetics and systematics, Baenidae, Baenodda, Chelydridae, Eileanchelys, Platysternon, Protostegidae

Jérémy Anquetin (2012). "Reassessment of the phylogenetic interrelationships of basal turtles (Testudinata)". Journal of Systematic Palaeontology 10 (1): 3–45. doi:10.1080/14772019.2011.558928.
Protostegidae

Anquetin, J., Barrett, P. M., Jones, M. E. H., Moore-Fay, S. & Evans, S. E., 2009: A new stem turtle from the Middle Jurassic of Scotland: new insights into the evolution and palaeoecology of basal turtles. Proceedings of the Royal Society of London: Biological series: Vol. 276, pp. 879-886 [doi: 10.1098/rspb.2008.1429] pdf.
Chelonian phylogenetics and systematics, Eileanchelys

Barley, A.J., Spinks, P.Q., Thomson, R.C., Shaffer, H.B.., 2010, Fourteen nuclear genes provide phylogenetic resolution for difficult nodes in the turtle tree of life. Molecular Phylogenetics and Evolution 55 (2010) 1189–1194 pdf
Morphology or Molecules?

Baur, George (1890). "On the Classification of the Testudinata". The American Naturalist 24 (282): 530–536. doi:10.1086/275138. JSTOR 2450882.
Protostegidae

Baur, G. 1891. Notes on some little known American fossil tortoises. Proc. Acad. Nat. Sci. Philadelphia, vol. 43, pp. 411-430.
Chelonian phylogenetics and systematics

Benton, M. J., Vertebrate Paleontology, 2nd ed. 2000, 3rd ed. 2004, Blackwell Science Ltd,
Australochelys, Casichelydia

Bhullar, B-A.S. and Bever, G.S. 2009. An Archosaur-Like Laterosphenoid in Early Turtles (Reptilia: Pantestudines). Breviora. vol.518:1-10. pdf
Mystery of Chelonian Origins

Gale A. Bishop. 2007. Handbook for SeaTurtle Interns especially on St. Catherines Island. part 5, Natural history of sea turtles pdf
Allopleuroninae, Dermochelyinae, Syllomus

Böhme, W. 2005. Handbuch der Reptilien und Amphibien Europas. Band 3/IIIB: Schildkröten (Testudines) II (Cheloniidae, Dermochelyidae, Fossile Schildkröten Europas) (Handbook of reptiles and amphibians of Europe, Volume 3/IIIB: Turtles (Testudines) II (Cheloniidae, Dermochelyidae, Fossil turtles of Europe)). Edited by Uwe Fritz. Illustrations: Igor G. Danilov and Jiri Moravec. Wiebeisheim: AULA-Verlag pdf (in German)

Borsuk-Bialynicka, M, & Evans, S.E. (2009). A long-necked archosauromorph from the Early Triassic of Poland. Paleontologica Polonica 65: 203–234. pdf
Mystery of Chelonian Origins

Boule, Marcellin. Priviteau, Jean. Les Fossiles. Éléments De Paléontologie. Libraries de L'académie Médecine. Paris, 1935. p. 433
Pleurosternon

Bour, R & Dubois, A. 1986, Nomenclature ordinale et familiale des tortues (Reptilia). Note complémentaire. Bulletin Mensuel de la Société Linnéenne de Lyon 55:87-90..
Classification, Meiolanoidea

Brian W. Bowen, 2003, What Is a Loggerhead Turtle? - The Genetic Perspective
Caretta, Cheloniinae

Donald B. Brinkman, 2009, A sea turtle skull (Cheloniidae: Carettini) from the Lower Miocene Nye formation of Oregon, U. S. A. Paludicola 7(2): 39-46 May 2009 pdf
Procolpochelys, Syllomus

Brochu, C.A. 2001. Progress and future directions in archosaur phylogenetics J. Paleont., 75(6), 2001, pp. 1185–1201 pdf
Mystery of Chelonian Origins

Edwin A. Cadena, Daniel T. Ksepka and Mark A. Norell (2013) New material of Mongolemys elegans Khosatzky and Mlynarski, 1971 (Testudines: Lindholmemydidae), from the Late Cretaceous of Mongolia with Comments on Bone Histology and Phylogeny. American Museum Novitates 3766 :1-28. 2013 doi: http://dx.doi.org/10.1206/3766.2 pdf
Mongolemys

Carroll, R. L., 1988: Vertebrate paleontology and evolution. W. H. Freeman and company, New York,
Chelonian phylogenetics and systematics

Cherepanov GO. 1997. The origin of the bony shell of turtles as a unique evolutionary model in reptiles. Russ J Herpet. 4:155–162
Mystery of Chelonian Origins

Ckhikvadze, Viacheslav M. 1971. First find of a Tertiary turtle of the family Platysternidae. Paleontal. Zhur., 1971, no. 4, pp. 137-139.
Macrocephalochelys

Colbert, E.H., 1969 Evolution of the Vertebrates, John Wiley & Sons, 2nd ed.
Chelonian phylogenetics and systematics

Collins, J. 1972. The chelonian Rhinochelys Seeley from the Upper Cretaceous of England and France. Palaeontolgy 13: 355 -378
Rhinochelys

Danilov, I.G.; Averianov, A.O; & Yarkov, A.A. (2010). Itilochelys rasstrigin gen. et sp. nov., a new hard-shelled sea turtle (Cheloniidae sensu lato) from the Lower Palaeocene of Volgograd Province, Russia. Proceedings of the Zoological Institute RSA 314 (1): 24–41.
Itilochelys

Danilov I.G., Claude J., Sukhanov V.B. A redescription of Elkemys australis (Yeh, 1974), a poorly known basal testudinoid turtle from the Paleocene of China // Proc. Zool. Inst. RAS. 2012. V. 316. ? 3. P. 223-238. pdf
Lindholmemydidae

Danilov, I. G. & Parham, J. F. 2006. A redescription of Plesiochelys tatsuensis from the Late Jurassic of China, with comments on the antiquity of the Extant Turtles Journal of Vertebrate Paleontology 28(2):306-318.
Morphology or Molecules?, Pantrionychia

Danilov I.G. and Parham J.F. 2008. A reassessment of some poorly known turtles from the Middle Jurassic of China, with comments on the antiquity of extant turtles.Journal of Vertebrate Paleontology, 28: 306–318
Morphology or Molecules? - abstract and text figures

deBraga, M & O Rieppel (1997) Reptile phylogeny and the interrelationships of turtles, Zool. J. Linnean Soc. 120: 281-354.
Mystery of Chelonian Origins

Dryden, L. S. 1988. Paraphyly of the Cryptodira and phylogenetic systematics of turtles. Unpublished Master's thesis, University of Kansas, Lawrence, KS.
Chelonian phylogenetics and systematics

Alain Dubois & Roger Bour, 2010 The distinction between family-series and class-series nomina in zoological nomenclature, with emphasis on the nomina created by Batsch (1788, 1789) and on the higher nomenclature of turtles, Bonn zoological Bulletin 57 (2): 149–171
Chelonii

Ehret, Dana Joseph 2004 "Skeletochronology as a method of aging Oligocene Gopherus laticuneus and Stylemys nebrascensis, using Gopherus polyphemus as a modern analog" Thesis, University of Florida. pdf
Hadrianus

Fong JJ, Brown JM, Fujita MK, Boussau B (2012) A Phylogenomic Approach to Vertebrate Phylogeny Supports a Turtle-Archosaur Affinity and a Possible Paraphyletic Lissamphibia. PLoS ONE 7(11): e48990. doi:10.1371/journal.pone.0048990 online

Fossil Turtle Newsletter January, 2004. Compiled by Walter Joyce and James Parham pdf

Fujita, M.K.; Tag N. Engstrom, David E. Starkey and H. Bradley Shaffer (2004). Turtle phylogeny: insights from a novel nuclear intron. Molecular Phylogenetics and Evolution 31 (3): 1031-1040
Trionychoidea

Benjamin P. Kear, and Michael S. Y. Lee, 2006, A primitive protostegid from Australia and early sea turtle evolution Biol. Lett. 2, 116-119 doi:10.1098/rsbl.2005.0406 pdf
Bouliachelys, Toxochelyidae

Krenz, James G.; Gavin J.P. Naylor; H. Bradley Shaffer and Fredric J. Janzen (2005) Molecular phylogenetics and evolution of turtles. Molecular Phylogenetics and Evolution, Volume 37 (1):178-191.
Morphology or Molecules?

R. Kohler (1995): A new species of the fossil turtle Psephophorus (Order Testudines) from the Eocene of the South Island, New Zealand, Journal of the Royal Society of New Zealand, 25:3, 371-384 pdf
Psephophorus

Gadagkar, S.R., Rosenberg, M.S., & Kumar, S. (2005). Inferring Species Phylogenies From Multiple Genes: Concatenated Sequence Tree Versus Consensus Gene Tree. Journal of Experimental Zoology (Mol Dev Evol) 304B:64–74 pdf
Morphology or Molecules?

Gaffney, E.S. 1972. The systematics of the North American family Baenidae (Reptilia Crytodira). Bull. Amer. Nat. Hist. 147(5):241-320.
Baenidae

Gaffney, E.S. (1975): A phylogeny and classification of higher categories of turtles. Bulletin of the American Museum of Natural History 155(5): 387-436. abstract, pdf
Testudinata/Testudines, Chelonian phylogenetics and systematics, Paracryptodira

Gaffney, Eugene S. (1975b). Phylogeny of the chelydrid turtles. Fieldiana, Geology (Chicago: Field Museum of Natural History) 33 (9): 166 (16). OCLC 1991971. online
Chelydrinae, Macrocephalochelys, Platysternidae, Platysternon

Gaffney, E.S. 1979. Comparative cranial morphology of Recent and fossil turtles. Bull. Amer. Mus. Natur. Hist. 164; 65-376
Testudinoidae

Gaffney, E.S (1984) Historical Analysis of Theories of Chelonian Relationship. Systematic Zoology 33. 3. 283-301.
Chelonian phylogenetics and systematics

Gaffney, E.S (1996) The postcranial morphology of Meiolania platyceps and a review of of the Meiolaniidae. Bulletin of the American Museum of Natural History ; no. 229 abstract and full pdf .
Chelydridae, Meiolania, Meiolaniidae, Ninjemys, Niolamia

Gaffney, E. S., Balouet, J. C., & Broin, F. (1984). New occurrences of extinct meiolaniid turtles in New Caledonia. American Museum novitates ; no. 2800 pdf
Meiolaniidae

Gaffney and Kitching 1994, The most ancient African turtle, Nature 369: 55-58, [doi:10.1038/369055a0]
Australochelys

Gaffney and Kitching 1995, The Morphology and Relationships of Australochelys, an Early Jurassic Turtle from South Africa, Am. Mus. Novitates 3130: 29 pp.
Australochelys

Gaffney, E. S., Kool, L., Brinkman, D. B., Rich, T. H. V., & Rich, Pat Vickers. 1998. Otwayemys, a new cryptodiran turtle from the early Cretaceous of Australia. American Museum novitates ; no. 3233 abstract & pdf
Otwayemys

Gaffney, ES & PA Meylan (1988), A phylogeny of turtles, in MJ Benton (ed.), The Phylogeny and Classification of the Tetrapods, 1: 157-219.
Pancheloniidae, Platysternon, Podocnemoidae, Testudinoidae, Trionychoidea, Xerobatinae

Gaffney E.S., Meylan, P. A., & Wyssz, A.R. 1991. A computer assisted analysis of the relationships of the higher categories of turtles. Cladistics 1991 7: 313-335 pdf
Chelonian phylogenetics and systematics, Chelydridae

Gauthier, J, AG Kluge & T Rowe (1988), The early evolution of the Amniota, in MJ Benton (ed.) The Phylogeny and Classification of the Tetrapods, Volume 1: Amphibians, Reptiles, Birds. Clarendon Press, pp. 103-155.
Testudines

Gauthier, J., D. C. Cannatella, K. De Queiroz, A. G. Kluge, & T. Rowe. 1989. Tetrapod phylogeny. In B. Fernholm, K. Bremer, and H. Jornvall (eds.) The Hierarchy of Life: 337-353. New York: Elsevier Science Publishers B. V. (Biomedical Division). pdf
Chelonian phylogenetics and systematics

J. Gerlach 2001 Tortoise phylogeny and the 'Geochelone' problem. Phelsuma, Volume 9 (supplement A) pdf
Megalochelyini, Xerobatinae

Gingerich, P. D., M. Haq, I. S. Zalmout, I. H. Khan, and M. S. Malkani. 2001. Origin of whales from early artiodactyls: hands and feet of Eocene Protocetidae from Pakistan. Science, 293: 2239-2242. pdf.
Mystery of Chelonian Origins

Jean-Michel Guillon, Loreleď Guéry, Vincent Hulin, & Marc Girondot, 2012, A large phylogeny of turtles (Testudines) using molecular data, Contributions to Zoology, 81 (3) 147-158
Megalochelyini, Xerobatinae

S. H. Haughton and A. S. Brink. 1954. A bibliographical list of Reptilia from the Karroo Beds of South Africa. Palaeontologia Africana 2:1-187.
Chelonian phylogenetics and systematics

Hay, 0. P. 1905. On the group of fossil turtles known as the Amphichelydia, with remarks on the origin and relationships of the suborders, superfamilies and families of Testudines. Bull. Amer. Mus.Nat. Hist., vol. 21, art. 9, pp. 137-175.
Chelonian phylogenetics and systematics

Hay, O.P. 1908. The fossil turtles of North America. Publications of the Carnegie Institution 75:1-568.
Chelonian phylogenetics and systematics, Archelon, Platysternon

Hedges, S. Blair, and Poling, Laura L. 1999, A Molecular Phylogeny of Reptiles. Science, Vol. 283, No.5404, pp.998-1001. pdf.
Mystery of Chelonian Origins

Hill, Robert V.. 2005. Integration of Morphological Data Sets for Phylogenetic Analysis of Amniota: The Importance of Integumentary Characters and Increased Taxonomic Sampling. Systematic Biology 54(4):530-547. abstract pdf.
Mystery of Chelonian Origins

Hirayama, R. 1984. Cladistic analysis of batagurine turtles (Batagurinae: Emydidae: Testudinoides); a preliminary result. Stud. Geol. Salamanticesia. 1; 141-157
Testudinoidae

Hirayama, Ren (1994). "Phylogenetic systematics of chelonioid sea turtles". Island Arc 3 (4): 270–284. doi:10.1111/j.1440-1738.1994.tb00116.x. Retrieved 2007-09-04.
Protostegidae, Rhinochelys

Hirayama, R. 1998. "Oldest known sea turtle". Nature 392 (6677): 705–708. doi:10.1038/33669
Protostegidae, Santanachelys

Hirayama, R. 2012 Review of fossil chelonioid sea turtles (Class Reptilia: Order Testudines: Chelonioidea) from Japan, Japan Geoscience Union Meeting 2012 pdf

Ren Hirayama, Donald B. Brinkman, and Igor G. Danilov, 2000, Distribution and Biogeography of Non-marine Cretaceous Turtles, Russian Journal of Herpetology Vol. 7, No. 3, 2000, pp. 181-198, pdf,
Plesiochelyidae, Pleurosternidae

Hirayama, R., Sakurai, K., Chitoku, T., Kawakami, G., and Kito, N. 2001. Anomalochelys angulata, an unusual land turtle of family Nanhsiungchelyidae (Superfamily Trionychoidea; Order Testudines) from the Upper Cretaceous of Hokkaido, North Japan. Russian Journal of Herpetology 8: 127-138.
Anomalochelys

Honegger, R. E. 1981. List of amphibians and reptiles either known or thought to have become extinct since 1600. Biological Conservation 19: 141-158.
Tortoise Resurrection

Hooks, III, G. E., 1998: Systematic revision of the Protostegidae, with a redescription of Calcarichelys gemma Zangerl, 1953. Journal of Vertebrate Paleontology: Vol. 18, #1, pp. 85-98 abstract
Protostegidae, Protosteginae

Hugall, A. F., Foster, R. & Lee, M. S. Y. 2007 Calibration choice, rate smoothing, and the pattern of tetrapod diversification according to the long nuclear gene RAG-1. Syst. Biol. 56, 543–563.
Mystery of Chelonian Origins

Hutchison, J. H. 1992. Western North American reptile and amphibian record across the Eocene/Oligocene boundary and its climatic implications. Prothero, D. R. & Berggren, W. A. (eds.), Eocene-Oligocene Climatic and Biotic Evolution 451-463.
Platysternon

Hutchison, J.H., 1996. Testudines. Cambridge University Press, Cambridge
Emydidae

Iverson J.B., Brown R.M., Akre T.M., Near T.J., Le M., Thomson R.C., Starkey D.E. 2007. In search of the tree of life of turtles.In: Shaffer H.B., FitzSimmons N.N., Georges A., Rhodin A.G.J., editors. Defining Turtle Diversity: Proceedings of a Workshop on Genetics, Ethics, and Taxonomy of Freshwater Turtles and Tortoises. Cambridge (MA). Lunenburg (MA): Chelonian Research Foundation. p. 85¨C106 (Chelonian research monographs).
Chelonian phylogenetics and systematics, Geoemydidae, Natator

Jackson, A.A.; 2008: Bedrock geology UK south. An explanation of the bedrock geology map of England and Wales - 1:625,000 edition, Keyworth, Nottingham, British Geological Survey,
Pleurosternon

Jamniczky, HA & Russell, AP (2007). Reappraisal of patterns of nonmarine cryptodiran turtle carotid circulation: evidence from osteological correlates and soft tissues. J Morphol. 2007 July; 268(7):571-87. pdf
Trionychoidae

Janke, A. D., Erpenbeck, D., Nilsson, M., and Arnason, U. 2001. The mitochondrial genomes of the iguana (Iguana iguana) and the caiman (Caiman crocodylus): implications for amniote phylogeny. Proc. R. Soc.B 268: 623-631.
Mystery of Chelonian Origins

Jehle, M. 2006, Paleocene mammals of the world - Turtles, Business as Usual
Adocus

Jones MEH, Werneburg I, Curtis N, Penrose R, O’Higgins P, et al. (2012) The Head and Neck Anatomy of Sea Turtles (Cryptodira: Chelonioidea) and Skull Shape in Testudines. PLoS ONE 7(11): e47852. doi:10.1371/journal.pone.0047852 online
Crown Chelonioidea

Joyce W.G. (2000) The first complete skeleton of Solnhofia parsonsi (Cryptodira, Eurysternidae) from the Upper Jurassic of Germany and its taxonomic implications J. Paleont., 74(4), 2000, pp. 684-700
pdf

Joyce W.G. (2007) Phylogenetic relationships of Mesozoic turtles. Bull. Peabody Mus Nat Hist 48(1):3–102
Chelonian phylogenetics and systematics, Baenidae, "Baenini",, Chelydridae, Pancheloniidae, Platysternon, Protostegidae

Joyce, WG, & Gauthier, JA, (2004), Palaeoecology of Triassic stem turtles sheds new light on turtle origins, Proc. R. Soc. Lond. B (2004) 271, 1–5 1 ĆÉ 2003 The Royal Society DOI 10.1098/rspb.2003.2523. pdf
Palaeochersis

Joyce, W.G., Jenkins, F.A., Jr., and Rowe, T. 2006. The presence of cleithra in the basal turtle Kayentachelys aprix; pp. 93-103 in Danilov, I.G. and Parham, J.F. (eds.), Fossil Turtle Research, Volume 1. Russian Journal of Herpetology 13 (suppl.). pdf
Mystery of Chelonian Origins

Joyce, W. G., Lucas, S. G., Scheyer, T. M., Heckert, A. B., and Hunt, A. P., 2008. A thin-shelled reptile from the Late Triassic of North America and the origin of the turtle shell Proceedings of the Royal Society B Published online (FirstCite). DOI: 10.1098/rspb.2008.1196. pdf.
Chelonian phylogenetics and systematics, Chinlechelys

Joyce, W. G. and Lyson, T. R., 2010, A neglected lineage of North American turtles fills a major gap in the fossil record. Palaeontology, 53: 241–248. doi: 10.1111/j.1475-4983.2009.00929.x abstract.
Plastomeninae

Joyce, W.G., Parham, J.F., & Gauthier, J.A., (2004), Developing a protocol for the conversion of rank-based taxon names to phylogenetically defined clade names, as exemplified by turtlesJ. Paleont., 78(5), 2004, pp. 989–1013. 0022-3360/04/0078-989$03.00 pdf.
Chelonian phylogenetics and systematics, Pancheloniidae, Pancheloniodea

Walter G. Joyce, Ariel Revan, Tyler R. Lyson and Igor G. Danilov (2009). Two New Plastomenine Softshell Turtles from the Paleocene of Montana and Wyoming. Bulletin of the Peabody Museum of Natural History 50 (2): 307–325. pdf
Plastomeninae

H. F. Kaddumi (2006). "A new genus and species of gigantic marine turtles (Chelonioidea: Cheloniidae) from the Maastrichtian of the Harrana Fauna-Jordan". Vertebrate Paleontology (journal) 3 (1): 1–14.
Gigantatypus

H.-V. Karl - Zur Taxonomie der känozoischen Weichschildkröten Österreichs und Deutschlands (Trionychidae: Trionychinae).- Mitt. Geol. Und Paläont. Landesmuseum Joanneum, 56: 273- 328; Graz (1998).
Trionyx messelianus

Karl, H.-V. (2002): Übersicht über die fossilen marinen Schildkrötenfamilien Zentraleuropas (Reptilia, Testudines). Mauritiana (Altenburg), 18 (2): 171-202.
Toxochelys

Karl, H.-V. (2005): The homology of supramarginals in turtles (Reptilia:Chelonii). Studia Geologica Salmanticensia, 41, pp. 63-75.
Mystery of Chelonian Origins

Hans-Volker Karl & Bent E. K. Lindow, 2009, First evidence of a Late Cretaceous marine turtle (Testudines: Chelonioidea) from Denmark Studia Geologica Salmanticensia, 45 (2): pp. 175-180
Toxochelys

S. A. Karl, B. W. Bowen, and J. C. Avise,1995, Hybridization among the ancient mariners: Identification and characterization of marine turtle hybrids with molecular genetic assays. Journal of Heredity 86:262-268.pdf
Cheloniinae

Killian JK, Buckley TR, Stewart N, Munday BL, Jirtle RL. 2001. Marsupials and Eutherians reunited: genetic evidence for the Theria hypothesis of mammalian evolution. Mamm. Genome 12:513–17 pdf
Mystery of Chelonian Origins

Kordikova, E.G. Comparative morphology of the palate dentition in Proganochelys quenstedti Baur 1887 from the Upper Triassic of Germany and chelonian ancestry. - N. Jb. Geol. Paläont. Abh., 225, 2: 195-249.
Proganochelys

Kullberg, M., Hallström, B. M., Arnason, U. and Janke, A. (2008), Phylogenetic analysis of 1.5 Mbp and platypus EST data refute the Marsupionta hypothesis and unequivocally support Monotremata as sister group to Marsupialia/Placentalia. Zoologica Scripta, 37: 115–127. doi: 10.1111/j.1463-6409.2007.00319.x
Mystery of Chelonian Origins

Kumazawa, Y. and Nishida, M. 1999. Complete Mitochondrial DNA Sequences of the Green Turtle and Blue-Tailed Mole Skink: Statistical Evidence for Archosaurian Affinity of Turtles. Mol. Biol. Evol. vol.16(6):784-792
Mystery of Chelonian Origins

Lambe, L. M. (1902). "New genera and species from the Belly River Series (Mid-Cretaceous)". Contributions to Canadian Palaeontology 3: 23–81.
Plesiobaena

Minh Le, Christopher J. Raxworthy, William P. McCord, & Lisa Mertz (2006), A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes. Molecular Phylogenetics and Evolution 450 (2):
Megalochelyini, Testudinidae, Testudininae, Testudinini

Lee, M.S.Y. 1993. The Origin of the Turtle Body Plan: Bridging a Famous Morphological Gap. Science. vol.261: 1716 1720.
Mystery of Chelonian Origins

Lee, MSY (1996), Correlated progression and the origin of turtles. Nature 379: 812-815.
Testudines

Lee, M.S.Y. 1997. Pareiasaur Phylogeny and the Origin of Turtles. Zoo. J. Linn. Soc. vol.120:197-280
Mystery of Chelonian Origins

Lee, M. S. Y. 1997a. Reptile relationships turn turtle. Nature, 389:245–246.
Mystery of Chelonian Origins

Lee, M. S. Y. 2001. Molecules, morphology, and the monophyly of diapsid reptiles. Contrib. Zool. 70: 121–138.
Mystery of Chelonian Origins

Li, Chun; Xiao-Chun Wu, Olivier Rieppel, Li-Ting Wang & Li-Jun Zhao (2008). "An ancestral turtle from the Late Triassic of southwestern China". Nature 456: 497-501. doi:10.1038/nature07533.
Odontochelys, Chelonian phylogenetics and systematics, Mystery of Chelonian Origins

Lipka, T. R., F. Therrien, D. B. Weishampel, H. A. Jamniczky, W.G. Joyce, M. W. Colbert, AND D. B. Brinkman. 2006. A new turtle from the Arundel Clay facies (Potomac Formation, Early Cretaceous) of Maryland, U.S.A. Journal of Vertebrate Paleontology, 20:300-307.
Arundelemys, Paracryptodira

Liu, L., Pearl, D.K., 2007. Species trees from gene trees: reconstructing Bayesian posterior distributions of a species phylogeny using estimated gene tree distributions. Syst. Biol. 56 (3), 504–514.
Morphology or Molecules?

Lucas, S.G. 2001, Chinese Fossil Vertebrates, Columbia University Press, New York
Chengyuchelys, Nanhsiungchelys

Lynch, S. C., and J. F. Parham. 2003. The first report of hard-shelled sea turtles (Cheloniidae sensu lato) from the Miocene of California, including a new species (Euclastes hutchisoni) with unusually plesiomorphic characters. PaleoBios 23:21-25.
Syllomus

Lyson, T.R., Bever, G.S., Bhullar, B-A.S.,Joyce, G.J., Gauthier, J.A. 2010. Transitional fossils and the origin of turtles. Biol. Lett. doi:10.1098/rsbl.2010.0371 pdf
Mystery of Chelonian Origins

Lyson, T. & Gilbert, S.F. (2009). Turtles all the way down: loggerheads at the root of the chelonian tree. Evolution & Development 11:2, 133-135 (2009) DOI: 10.1111/j.1525-142X.2009.00325.x pdf
Mystery of Chelonian Origins

Tyler R. Lyson and Walter G. Joyce (2009). "A Revision of Plesiobaena (Testudines: Baenidae) and an Assessment of Baenid Ecology Across the K/T Boundary". Journal of Paleontology 83 (6): 833–853. doi:10.1666/09-035.1.
Baenidae, Plesiobaena

Tyler R. Lyson and Walter G. Joyce (2011). Cranial anatomy and phylogenetic placement of the enigmatic turtle Compsemys victa Leidy, 1856 Journal of Paleontology, 85(4), 2011, p. 789–801
Compsemys

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Protostegidae

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Natator

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Nanhsiungchelyidae

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Protostegidae

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Sandownia

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Hesperotestudo, Stylemys, Xerobatinae

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Pleurosternon

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Testudinidae, Toxochelys

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Mystery of Chelonian Origins

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Odontochelys

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Pleurosternon

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Pleurosternon

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Macrobaenidae

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Judithemys

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Morphology or Molecules?

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Caretta, Euclastes, Mexichelys, Pancheloniidae, Syllomus

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Megalochelyini

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Chengyuchelys, Xinjiangchelyidae

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Pleurosternon

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Macrocephalochelys

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Tortoise Resurrection

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Overview, Macrobaenidae, Sinemydidae

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Sinemydids and Macrobaenids, Xinjiangchelyidae

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Pleurosternon

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Chelonian phylogenetics and systematics

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Palaeochersis

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Morphology or Molecules?, Trionychoidea

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Meiolaniidae, Trionychoidea, Testudines.

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Mystery of Chelonian Origins

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Protostegidae

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Protostegidae

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Chelonian phylogenetics and systematics

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Hispaniachelys

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Caretta

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Chelonian phylogenetics and systematics, Condorchelys

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Chelonian phylogenetics and systematics, Morphology or Molecules?, Pleurodira, Protostegidae

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Kallokibotion

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Protostegidae

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Mongolochelys

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Mongolochelys

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Adocus

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Morphology or Molecules?, Xerobatinae

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Argillochelys

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Rhinochelys

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Archelon

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Archelon, Protostega

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Mystery of Chelonian Origins

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Calcarichelys, Chelosphargis, Protostega, Rhinochelys

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Toxochelyidae, Toxochelys

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Mystery of Chelonian Origins

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Mystery of Chelonian Origins

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Adocidae, Trionychoidea

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Manchurochelys

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Allopleuron, Argillochelys, Caretta, Chelonia, Chitracephalus, Corsochelys, Dermochelyinae, Dermochelys, Eochelone , Eosphargis, Euclastes, Hadrianus, Plesiochelyidae, Protosphargis, Psephophorus, Stylemys, Toxochelys

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Caretta




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