Palaeos Palaeos Chelonii
Vertebrates Dendrogram

Chelonii: Dendrogram

Abbreviated Dendrogram
ANAPSIDA ?  
|
o CHELONII
|--Odontochelys   
`--+--Proganochelys
   `--+--Proterochersis 
      `--+--Meiolanoidea
         `--+--Pleurodira
            |--Paracryptodira 
            `--+--Protostegidae
               `--+==Xinjiangchelyidae
                  `--+==Sinemydidae
                     |--Trionychoidea
                     `--+--+--Chelonioidea
                         |  |--Chelydridae
                         |  `--Kinosternoidea
                         `--Testudinoidea
Contents

Overview
Chelonian origins
Testudines
Casichelydia
Derived Stem Testudines
Meiolanoidea
Pleurodira
Paracryptodira
Eucryptodira
Protostegidae
Centrocryptodira
Trionychoidea
Chelonioidea
Chelydridae
Trionychoidea
Kinosternoidea
Testudinoidea
Testudinidae
Dendrogram
References







                   `--Polycryptodira MH
ANAPSIDA ?  WP, MH
|
o CHELONII ToL
|--Odontochelys 
`--o Testudinata
   |--Proganochelys ToL
   `--+--Chinlechelys
   `--o <>Casichelydia 
      |--Palaeochersis
      |--Australochelys 
      `--+--Proterochersis 
         `--o Cryptodira MH
            |--Kayentachelys •X ToL
            `--+--Condorchelys 
               |--Eileanchelys 
               `--+--Solemydidae
                  |--o Meiolanoidea (Joyce 2007;  Anquetin, 2009)
                  |  |--Chubutemys
                  |  |--Naomichelys
                  |  |--Chelycarapookus
                  |  `--+--+--Mongolochelys
                  |     |   `?--Otwayemys (if statistical morphology)
                  |     `--o Meiolaniidae
                  |        |--Niolamia
                  |        `--+--Ninjemys
                  |           `--Meiolania
                  `--+--Kallokibotion
                     |?-- Hispaniachelys
                     `--o Testudines crown group
                        |--o Pleurodira ToL, MH
                        |  |--Proterochersis •X ToL
                        |  `--o Eupleurodira
                        |     |--Chelidae MH, ToL
                        |     `--o Pelomedusoidea
                        |        |--o Podocnemoidae
                        |        |  |--Bothremyidae •XToL, MH
                        |        |  `--Podocnemidae MH
                        |        `--Pelomedusidae MH, ToL
                        |--o Paracryptodira 
                        |  |--Compsemys 
                        |  |--+-Dorsetochelys 
                        |  |  `--o Pleurosternidae 
                        |  |     |--Pleurosternon 
                        |  |     `--Glyptops 
                        |  `--+--Arundelemys 
                        |     `--o Baenidae 
                        |        |--Neurankylus 
                        |        `--o Baenodda 
                        |           |--o Plesiobaena-Palatobaena lineage 
                        |           |  |--Plesiobaena 
                        |           |  `--Palatobaena 
                        |           `--o Eubaena-Stygiochelys lineage 
                        |              |--+--Boremys 
                        |              |  `--Eubaena 
                        |              `--+--Stygiochelys 
                        |                 `--+--Baena 
                        |                    `--Chisternon 
                        `--o Eucryptodira

                `--+--o Paracryptodira 
                   |--o Pleurodira (Joyce 2007)
                   `--o Eucryptodira
                      |--+--Thalassemyidae
                      |  `--Protostegidae
                      `--+--Plesiochelyidae
                         `--+--Angolachelonia
                            `--+--Xinjiangchelyidae
                               `--+--o Pleurodira (Anquetin, 2009)
                                  `--o Centrocryptodira
                                     |--Meiolaniidae (Gaffney, 1996)
                                     `--+==Sinemydidae
                                              `--+?--Trionychoidea (molecular + stratigraphy)
                                           |== Macrobaenidae
                                           `--Durocryptodira
`--o Eucryptodira |--Thalassemyidae |--Protostegidae `--+--Plesiochelyidae `--+--o Angolachelonia | |--Solnhofia | `--+--Sandownia | `--Angolachelys `--+==Xinjiangchelyidae `--o Centrocryptodira |--Pleurodira `--o Eucryptodira ? |--Thalassemyidae | |--Protostegidae | |--Plesiochelyidae | `--+--o Angolachelonia | | |--Solnhofia | | `--+--Sandownia | | `--Angolachelys `---- ?--+--Chengyuchelyidae | |--Chengyuchelys | `--Xinjiangchelyidae `--o Centrocryptodira |?--Meiolaniidae (hennigian morphology) `--o Cryptodira crown group (molecular) |?--Trionychoidea (molecular + stratigraphy) `--o Durocryptodira (molecular + stratigraphy) |==Sinemydidae `--o Cryptodira crown group (morphology) |--Chelonioidea `--+--Chelydridae |--+?--Trionychoidea (morphology) | `--Kinosternoidea `--Testudinoidea `--o Centrocryptodira |---------------? (if molecular + stratigraphy) | `,--o Trionychoidae | | |--Basilochelys | | `--+--o Adocusia (stratigraphy implies paraphyly) | | | |--Yehguia | | | `--+--o Adocidae | | | | |-- Adocus | | | | `-- Shachemydinae | | | |--o Nanhsiungchelyidae | | | |--Kharakhutulia | | | `--+--Hanbogdemys | | | |--o "Nanhsiungchelyini" | | | | |--Anomalochelys | | | | `--Nanhsiungchelys | | | |--Zangerlia | | | `--Basilemys | | `--+--Peltochelys | | `--+--Crown group Trionychoidae | | |--Kizylkumemys | | `--+--Carettochelyidae | | `--o Trionychidae | | |--+--Plastomeninae | | | `--Cyclanorbinae | | `--Trionychinae |--+--o Trionychoidea | |--Basilochelys | `--+--o Adocusia | | |--Yehguia | | `--+--o Adocidae | | | |-- Adocus | | | `-- Shachemydinae | | `--o Nanhsiungchelyidae | | |--Kharakhutulia | | `--+--Hanbogdemys | | |--o "Nanhsiungchelyinae" | | | |--Anomalochelys | | | `--Nanhsiungchelys | | |--Zangerlia | | `--Basilemys | `--+--Peltochelys | `--o Trionychia | |--Carettochelyidae | `--o Trionychidae | |--+--Plastomeninae | | `--Cyclanorbinae | `--Trionychinae o Chelonioidea |--Chitracephalus `--+--Mesodermochelys `--+--Toxochelyidae `--+--Chelonioidea |--+--Euclastes | |--Puppigerus | `--Cheloniidae `--Dermochelyidae `--+--+--Chelonioidea | `--+--Chelydridae | |--Emarginachelys | `--o Kinosternoidea | |--Dermatemydidae | `--o Kinosternia | |--Agomphus | |--Hoplochelys | `--o Kinosternidae | |--Kinosterninae | `--Staurotypinae `--Testudinoidea `--o Centrocryptodira |--Trionychoidea |--+--Hoyasemys | `--Sinemydidae |?--Osteopygis `--+--Kirgizemys `--+--o Macrobaenidae | |--Judithemys | `--Macrobaena |--o Chelonioidea-Chelydridae-Kinosternoidea Clade `--+--Angolachelonia `--o Xinjiangchelyid - Crown Cryptodire clade |--Chengyuchelys |--Larachelus |==Xinjiangchelyidae `--o Centrocryptodira |==Sinemydidae (in part) |--Trionychoidea |--+--Chelonioidea | `--+--Emarginachelys | |--o Chelydroidea | | |--o Platysternidae | | | |--Macrocephalochelys | | | `--Platysternon | | `--o Chelydridae | | |--Protochelydra | | `--Chelydrinae | `--Kinosternoidea `--Testudinoidea `--+--+--Chelonioidea | `--+--Chelydridae | `--Kinosternoidea |--Trionychoidea `--+--+--Chelonioidea | `--+--Chelydridae | `--Kinosternoidea `--o Testudinoidea |==Lindholmemydidae incl. Mongolemys `--+-Emydidae `--o Testudinoidae |--Echmatemys `--+--Geoemydidae `--Testudinidae `--o Testudinidae |--Hadrianus `--+--o Xerobatinae | |--Manouria | `--+--Stylemys | `--+--Gopherus | `--Hesperotestudo `--o Testudininae |--Testudinini `--Megalochelyini

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