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Lycophyta after Cleal

Class Lycopsida ("club-mosses') Order Drepanophycales Family Drepanophycaceae Order Protolepidodendrales Family Protolepidodendraceae Family Eleutherophyllaceae Order Lycopodiales Family Lycopodiaceae Order Selaginellales Family Selaginellaceae Order Lepidocarpales Family Cyclostigmaceae Family Flemingitaceae Family Sigillariostrobaceae Family Lepidocarpaceae Family Spenceritaceae Family Caudatocarpaceae Family Pinakodendraceae Family Sporangiostrobaceae Family Pleuromeiaceae Order Miadesmiales Family Miadesmiaceae Order Isoetales ("quillworts') Family Isoetaceae Family Chaloneriaceae

Palaeozoic Palaeobotany of Great Britain as well as listing British sites and the history of discovery of plant fossils in Britian, this is an excellent intro to Palaeobotany in general.

C.J. Cleal, 1993, in The Fossil Record 2, Chapman & Hall, London

Lycophyta cladogram (various sources)

■──○Lycophyta ├─┬─○†Zosterophyllopsida │ │ ├─†Zosterophyllum │ │ ├─†Gosslingia │ │ ├─†Rebuchia │ │ ├─†Bathurstia │ │ ├─†Crenaticaulis │ │ ├─†Serrulacaulis │ │ ├─○†Sawdonia │ │ │ ├─†S. ornata │ │ │ └─†S. acanthotheca │ │ ├─†Discalis longistipa │ │ ├─†Drepanophycus spinaeformis │ │ ├─†Kaulangiophyton akantha │ │ └─†Asteroxylon mackiei │ └─┬─○†ASTEROXYLALES │ └─┬─○†Drepanophycales │ └─┬?─†Baragwanathia longifolia │ ├?-†Archaeosigillaria │ └─○LYCOPODIOPSIDA) │ └─○Lycopodiales │ └─○Lycopodiaceae │ ├─○Huperzia │ │ └─H. selago │ ├─○Lycopodiella │ │ └─L. inundata │ ├─○Lycopodium │ │ ├─L. annotinum │ │ └─L. clavatum │ └─○Diphasiastrum │ ├─D. alpinum │ └─D. complanatum └─┬─○†Protolepidodendrales │ ├─○†Prolepidodendron │ │ └─†P. scharyanum │ ├─○†Estinnophyton │ │ ├─†E. wahnbachense │ │ └─†E. gracilis │ ├─†Sugambrophyton │ ├─†Colpodexylon │ ├─†Leclercqia complexa │ └─○†Spencerites │ ├─†S. moorei │ └─†S. insignis └─┬?─○Selaginellales │ └─○Selaginellaceae │ ├─†Selagibellites hallei │ ├─†Miadesmia │ └─○Selaginella │ ├─†S. harrisiana │ └─S. selaginoides └─○Isoetopsida ├─○Lepidodendrales │ ├─†Valmeyerodendron triangularifolium │ ├─†Cyclostigma │ ├─†Bisporangiostrobus │ ├─†Lepidosigillaria │ ├─○†Sigillariaceae │ │ └─○†Sigillaria │ │ ├─†(Eusigillaria) │ │ ├─†(Subsigillaria) │ │ └─†(Sigillaria) │ └─○†Lepidodendraceae │ ├─○†Lepidodendron │ │ ├─†L. hickii │ │ └─†L. aculaetum │ ├─†Paralycopodites brevifolius │ ├─○†Diaphorodendron │ │ ├─†D. vaskulare │ │ ├─†D. phillipsii │ │ ├─†D. scleroticum │ │ └─†D. dicentricum │ └─○†Lepidophloios │ └─†L. halli └─○Isoetales ├─○Isoetaceae [Isoëtaceae] │ └─○Isoetes [Isoëtes] │ ├─I. lakustris │ └─I. echinospora ├─○Pleuromeiales │ ├─†Pleuromeia sternbergi │ ├─†Nathorstiana arborea │ └─†Chaloneria ├─†Paurodentron fraipontii ├─†Oxroadia └─†Prostigmaria eggertiana

Reference(s):

Hämet-Ahti, L., Suominen, J., Ulvinen, T., Uotila, P. & Vuokko, S., 1984: Retkeilykasvio. --Suomen Luonnonsuojelun Tuki Oy & Forssan Kirjapaino Oy, Forssa, 1984, 1-544

--iNet: The Tree of Life

Steward, W. N. & Rothwell, G. W., 1993: Paleobotany and the Evolution of Pants, 2nd Edition. --Cambridge University Press, Cambridge, U.S.A., xii-521

Links

Systematics of the Lycophyta, at the UCMP site has a simplified and incomplete lycopsid phylogeny -- but useful for getting oriented to the main groups. Another fine introduction, which will probably not be on the web very long, can be found at An Overview of Green Plant Phylogeny. A nice student paper with an interesting phylogeny: Brautigam, S, C Li, A Mudry, G Miller & E Prebil (2001), An analysis of the evolution of the vascular cryptogams, Lycopodophyta, Sphenophyta, and the Pteridophyta, through examination of extinct and extant genera. J. Sys. Biol. Susq. Univ. 8(4). See Brautigam et al. (2001). The ultimate collection of lycopsid links in general is, as usual, on the Links for Paleobotanists site. See Lycophyta, Introductions to both Fossil and Recent Plant Taxa.  Having little to do with lycopsid phylogeny per se, but well worth reading: Bateman, RM, PR Crane, WA DiMichele, PR Kenrick, NP Rowe, T Speck & WE Stein (1998), Early evolution of land plants: Phylogeny, physiology, and ecology of the primary terrestrial radiation. Ann. Rev. Ecol. Syst. 29: 263–92 See Bateman et al. (1998). ATW050803.

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page uploaded on Kheper site 14 November 1999, on this site 31 March 2002
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dendrogram © Mikko Haaramo
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