Palaeos Palaeos Sauropterygia
Vertebrates Dendrogram

Sauropterygia: Dendrogram

Abbreviated Dendrogram
DIAPSIDA
|--ARCHOSAUROMORPHA
|LEPIDOSAUROMORPHA
|--SAUROPTERYGIA
|  |--Placodontia
|  |==Pachypleurosauridae
|  `--+--Nothosauridae
|     `--Plesiosauria
|        |--Pliosauroidea
|        `--Plesiosauroidea
|           |--Elasmosauridae
|           `--Cryptocleidoidea
`--Lepidosauriformes
   |--SPHENODONTIA
   `--SQUAMATA            
Contents

Overview
Sauropterygia
Placodontia
Pachypleurosauridae
Nothosauridae
Pistosauridae
Plesiosauria
Pliosauroidea
Plesiosauroidea
Elasmosauridae
Cryptocleididea
Polycotylidae
Dendrogram
References


DIAPSIDA
|--ARCHOSAUROMORPHA
|
LEPIDOSAUROMORPHA ToL
|--SAUROPTERYGIA X (placodonts + plesiosaurs) MH
|  |--Placodontia •X MH HCW08
|  |   |--Paraplacodus •X
|  |   `--+--Placodus •X
|  |       `--Cyamodontoidea •X
|  |           |--+--Cyamodus •X
|  |           |  `--Henodus •X
|  |           `--Placochelyidae •X
|  |               |--Placochelys •X
|  |               `--Psephoderma •X
|  `--Nothosauria X (plesiosaurs > placodonts)
|     |--Wumengosaurus
|     |--Pachypleurosauridae X MH Rieppel (1998), HCW08
|     |  |--Hanosaurus •X    
|     |  `--+--+--Dactylosaurus •X 
|     |     |  `--Anarosaurus •X  
|     |     `--+--Serpianosaurus •X  S89--Neusticosaurus •X (O'K&S99)
|     `--Eusauropterygia X (nothosaurs + plesiosaurs)
|        |--Nothosauridae X (nothosaurs > plesiosaurs) MH  
|        |  |--Keichousaurus •X HCW08 
|        |  `--+--Simosaurus •X HCW08 
|        |     `--+--Nothosaurus •X HCW08
|        |        `--+--Ceresiosaurus •X
|        |           `--Lariosaurus •X HCW08
|        `--Pistosauroidea X   (plesiosaurs > nothosaurs)
|           |--Corosaurus •X HCW08
|           `--+--Cymatosaurus •X  R97
|              `--+--Pistosaurus •X HCW08
|                  `--+--Yunguisaurus
|                     `--+--Augustasaurus
|                        `--Plesiosauria X MH
|                           |--Pliosauroidea X MH
|                           |  |--Bishanopliosaurus •X
|                           |  |--Thalassiodracon •X  
|                           |  `--+--Archaeonectrus •X
|                           |     |--Eurycleidus •X
|                           |     `--+--Rhomaleosauridae X MH
|                           |        |  |--Yuzhoupliosaurus •X  
|                           |        |  `--+--Rhomaleosaurus •X  
|                           |        |     `--+--Simolestes •X
|                           |        |        `--Leptocleidus •X
|                           |        `--Pliosauridae X MH
|                           |           |--Maresaurus •X  
|                           |           |--Macroplata •X  
|                           |           `--+--Kronosaurus •X
|                           |              `--+--Peloneustes •X  
|                           |                 `--+--Liopleurodon •X  
|                           |                    `--+--Pliosaurus •X 
|                           |                       `--Brachauchenius •X
|                           `--Plesiosauroidea X
|                              |--Attenborosaurus •X  
|                              `--+--Plesiosaurus •X
|                                 `--+--Eretmosaurus •X  
|                                    |--Sthenarosaurus •X
|                                    `--+--Microcleidus •X
|                                       `--+--Elasmosauridae •X MH
|                                          `--Cryptocleidoidea X MH
|                                             |--Cryptocleididae X
|                                             |  |--Colymbosaurus •X
|                                             |  |--Cryptocleidus •X
|                                             |  `--Muraenosaurus •X
|                                             `--Tricleidia X
|                                                |--Tricleidus •X
|                                                `--+--Cimoliasauridae X
|                                                   |  |--Kimmerosaurus •X  
|                                                   |  `--Cimoliasaurus •X
|                                                   `--Polycotylidae •X
|                                                        |--Plesiopleurodon •X
|                                                        `--+--Dolichorhynchops •X
|                                                            `--Trinacromerum •X
`--Lepidosauriformes MH
   |--Kuehneosauridae •X
   `--Lepidosauria
      |--SPHENODONTIA
      `--SQUAMATA

Alternate dendrogram, taking into account bone histology studies by Klein 2010, Hugi et al 2011, and Hugi 2011. Taxa for which there is information are in bold. The status of many taxa is uncertain due to lack of histological data:

SAUROPTERYGIA X 
   |?--Hanosaurus •X  
   |--+--Serpianosaurus •X --Neusticosaurus •X 
   |  `--Nothosauridae X (nothosaurs > plesiosaurs) 
   |      |--Keichousaurus •X 
   |      `--+--Simosaurus •X 
   |           `--+--Nothosaurus •X 
   |              `--+--Ceresiosaurus •X
   |                 `--Lariosaurus •X 
    `--+--+--+--Dactylosaurus •X 
       |    |   `--Anarosaurus •X  
       |    `--Placodontia •X 
       |        |--Paraplacodus •X
       |        `--+--Placodus •X
       |            `--Cyamodontoidea •X
       |               |--+--Cyamodus •X
       |               |  `--Henodus •X
       |               `--Placochelyidae •X
       |                   |--Placochelys •X
       |                   `--Psephoderma •X
       `--Pistosauroidea X   (plesiosaurs > nothosaurs)
           |--Corosaurus •X 
           `--+--Cymatosaurus •X 
              `--+--Pistosaurus •X 
                  `--+--Yunguisaurus
                     `--+--Augustasaurus
                        `--Plesiosauria X 

Here the placodont-nothosauria, and pachypleurosaur-eusauropterygia clades are replaced by a nothosaur-pistosaur topology, with the pachypleurosaurs spread among both (and perhaps also basal to botrh?) and the placodonts included with the pistosaurs. As with molecular-based studies, this gives a very different result to the conventional morphology-based analysis. MAK111102




last revised ATW080330, MAK111102

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