Palaeos Palaeos Lycopodiophyta
Plants Classification &
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Kingdoms Index Systematics

 

Lycophyta after Cleal

    Class Lycopsida ("club-mosses')
        Order Drepanophycales
            Family Drepanophycaceae
        Order Protolepidodendrales
            Family Protolepidodendraceae
            Family Eleutherophyllaceae
        Order Lycopodiales
            Family Lycopodiaceae
        Order Selaginellales
            Family Selaginellaceae
        Order Lepidocarpales
            Family Cyclostigmaceae
            Family Flemingitaceae
            Family Sigillariostrobaceae
            Family Lepidocarpaceae
            Family Spenceritaceae    
            Family Caudatocarpaceae
            Family Pinakodendraceae
            Family Sporangiostrobaceae
            Family Pleuromeiaceae        
        Order Miadesmiales 
            Family Miadesmiaceae
        Order Isoetales ("quillworts')
            Family Isoetaceae
            Family Chaloneriaceae 

web page Palaeozoic Palaeobotany of Great Britain as well as listing British sites and the history of discovery of plant fossils in Britian, this is an excellent intro to Palaeobotany in general.

C.J. Cleal, 1993, in The Fossil Record 2, Chapman & Hall, London


Lycophyta cladogram (various sources)

<==o LYCOPHYTA 
   |--+--o †ZOSTEROPHYLLOPSIDA
   |  |  |-- †Zosterophyllum
   |  |  |-- †Gosslingia
   |  |  |-- †Rebuchia
   |  |  |-- †Bathurstia
   |  |  |-- †Crenaticaulis
   |  |  |-- †Serrulacaulis
   |  |  |--o †Sawdonia
   |  |  |  |-- †S. ornata
   |  |  |  `-- †S. acanthotheca
   |  |  |-- †Discalis longistipa
   |  |  |-- †Drepanophycus spinaeformis
   |  |  |-- †Kaulangiophyton akantha
   |  |  `-- †Asteroxylon mackiei
   |  `--+--o †ASTEROXYLALES
   |     `--+--o †DREPANOPHYCALES
   |        `--+?- †Baragwanathia longifolia
   |           |?- †Archaeosigillaria
   |           `--o LYCOPODIOPSIDA)
   |              `--o LYCOPODIALES
   |                 `--o Lycopodiaceae 
   |                    |--o Huperzia 
   |                    |  `-- H. selago
   |                    |--o Lycopodiella
   |                    |  `-- L. inundata 
   |                    |--o Lycopodium 
   |                    |  |-- L. annotinum
   |                    |  `-- L. clavatum 
   |                    `--o Diphasiastrum
   |                       |-- D. alpinum 
   |                       `-- D. complanatum
   `--+--o †PROTOLEPIDODENDRALES
      |  |--o †Prolepidodendron
      |  |  `-- †P. scharyanum
      |  |--o †Estinnophyton 
      |  |  |-- †E. wahnbachense
      |  |  `-- †E. gracilis
      |  |-- †Sugambrophyton
      |  |-- †Colpodexylon
      |  |-- †Leclercqia complexa
      |  `--o †Spencerites 
      |     |-- †S. moorei
      |     `-- †S. insignis
      `--+?-o SELAGINELLALES 
         |  `--o Selaginellaceae 
         |     |-- †Selagibellites hallei
         |     |-- †Miadesmia
         |     `--o Selaginella 
         |        |-- †S. harrisiana
         |        `-- S. selaginoides 
         `--o ISOETOPSIDA 
            |--o LEPIDODENDRALES 
            |  |-- †Valmeyerodendron triangularifolium
            |  |-- †Cyclostigma
            |  |-- †Bisporangiostrobus
            |  |-- †Lepidosigillaria
            |  |--o †Sigillariaceae
            |  |  `--o †Sigillaria
            |  |     |-- †(Eusigillaria)
            |  |     |-- †(Subsigillaria)
            |  |     `-- †(Sigillaria)
            |  `--o †Lepidodendraceae
            |     |--o †Lepidodendron
            |     |  |-- †L. hickii
            |     |  `-- †L. aculaetum
            |     |-- †Paralycopodites brevifolius
            |     |--o †Diaphorodendron
            |     |  |-- †D. vaskulare
            |     |  |-- †D. phillipsii
            |     |  |-- †D. scleroticum
            |     |  `-- †D. dicentricum
            |     `--o †Lepidophloios
            |        `-- †L. halli
            `--o ISOETALES 
               |--o Isoetaceae [Isoλtaceae] 
               |  `--o Isoetes [Isoλtes] 
               |     |-- I. lakustris
               |     `-- I. echinospora
               |--o Pleuromeiales
               |  |-- †Pleuromeia sternbergi
               |  |-- †Nathorstiana arborea
               |  `-- †Chaloneria
               |-- †Paurodentron fraipontii
               |-- †Oxroadia
               `-- †Prostigmaria eggertiana

Reference(s):

Hδmet-Ahti, L., Suominen, J., Ulvinen, T., Uotila, P. & Vuokko, S., 1984: Retkeilykasvio. --Suomen Luonnonsuojelun Tuki Oy & Forssan Kirjapaino Oy, Forssa, 1984, 1-544

--iNet: The Tree of Life

Steward, W. N. & Rothwell, G. W., 1993: Paleobotany and the Evolution of Pants, 2nd Edition. --Cambridge University Press, Cambridge, U.S.A., xii-521


Links

Systematics of the Lycophyta, at the UCMP site has a simplified and incomplete lycopsid phylogeny -- but useful for getting oriented to the main groups.  Another fine introduction, which will probably not be on the web very long, can be found at An Overview of Green Plant Phylogeny.  A nice student paper with an interesting phylogeny: Brautigam, S, C Li, A Mudry, G Miller & E Prebil (2001), An analysis of the evolution of the vascular cryptogams, Lycopodophyta, Sphenophyta, and the Pteridophyta, through examination of extinct and extant generaJ. Sys. Biol. Susq. Univ. 8(4).  See Brautigam et al. (2001). The ultimate collection of lycopsid links in general is, as usual, on the Links for Paleobotanists site.  See Lycophyta, Introductions to both Fossil and Recent Plant Taxa.  Having little to do with lycopsid phylogeny per se, but well worth reading: Bateman, RM, PR Crane, WA DiMichele, PR Kenrick, NP Rowe, T Speck & WE Stein (1998), Early evolution of land plants: Phylogeny, physiology, and ecology of the primary terrestrial radiationAnn. Rev. Ecol. Syst. 29: 263–92 See Bateman et al. (1998).  ATW050803.


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page uploaded on Kheper site 14 November 1999, on this site 31 March 2002
last modified ATW040227
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dendrogram © Mikko Haaramo
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