Palaeos: Paleozoic Palaeos Cisuralian Epoch
Permian Period Artinskian Age

The Artinskian Age

The Artinskian Age of the Cisuralian Epoch: 284 to 276 million years ago



Introduction

Permian scene

As the climate became drier, the early Permian semiaquatic and lowland floodplain Dimetrodon and Eryops dominated fauna shown here, which had flourished for some 25 million years, was replaced by the more advanced floodplain and upland caesid and therapsid fauna.

An Artinskian bestiary: Edaphosaurid-Nectridean Province


Illustration by Robert Bakker

Early Permian tetrapod provinces

This map shows the three known tetrapod provinces (shaded) of the early Permian -
the tropical Edaphosaurid (centre),
the Kazakhstan Seymouriamorph province (upper right),
and the high latitude Gondwana Mesosaurid province (bottom)
From A.R. Milner, "Biogeography of Palaeozoic Tetrapods" fig.13.3; in J.A. Long (ed.) Palaeozoic Vertebrate Biostratigraphy and Biogeography, 1993, John Hopkins University Press, Baltimore

Known Occurrence of early Permian Tetrapod Faunas

Note: the following is based on Anderson and Cruikshank 1978 slightly modified.

Location tetrapod zone approx time *
USA
Western Europe
Eastern Europe
Brazil southern Africa India
Age     Texas Oklahoma Germany Russian platform Parana Basin South Africa / South-West Africa Himalayas
biotic province     Edaphosaurid province 
(equatorial Euramerica)
    Mesosaurid province
(south-west Gondwanaland)
Baigendzhinian 10 277

278

279
Choza Hennesey          
9 Vale          
8 Arroyo Garber   zone 0      
Aktasinian 7 280

281

282

283
Ludens West Grandfield          
6 Clyde Deep Red Run     Irati White band  
5 Belle Plains Wellington formation          

* approximate time in MYA (millions of years ago) - nearest million year intervals

Xenacanthus
Class Chondrichthyes
Subclass Elasmobranchii
Order Xenacanthida
Family Xenacanthidae

Xenacanthus


Length (total): about 1 to 2 meters
Diet: aquatic vertebrates
Comments:

Some Tetrapods - Aktasinian (Early Artinskian)



Class Tetrapoda
Order Temnospondyli
Superfamily Eryopoidea
Family Eryopidae

Eryops megacephalus [Cope]

Horizon: Wichita Series (Wolfcampian age), Texas Red Beds
Locality:
Specimens:
Length (skull):
Length (total): 1.5 to 2 meters
Weight: about 130 kg
Diet: other tetrapods
Comments: a large long-lived aquatic temnospondyl

Arroyo Formation - Early Baigendzhinian Age


Plesion Synapsida (Theropsida)
Order Pelycosauria
Suborder Eupelycosauria
Family Sphenacodontidae

Dimetrodon grandis (Case, 1907)

Horizon and Locality: Arroyo Formation, Clear Fork Group, Texas, USA
Specimens:
Length (skull): 42 cm
Length (total): 3.2 metres
Weight: around 250 kg
Diet: other tetrapods
Comments: The last and largest of the long-skulled, stocky-bodied lineage of dimetrodonts.  Apart from D. angelensis this was the largest species of Dimetrodon. A large, heavily-built, very common species, it is close to the earlier D. limbatus, with which it differs in larger size, more elongate and stouter neural spines, and fewer premaxillary teeth (only two on either side, the fewest for any species of the genus). It is associated with the slightly smaller but equally successful Dimetrodon gigashomogenes
References: Romer and Price 1940, Reisz 1986
Links:

Plesion Synapsida (Theropsida)
Order Pelycosauria
Suborder Eupelycosauria
Family Sphenacodontidae

Dimetrodon gigashomogenes Case, 1907

Horizon and Locality: Arroyo, Vale and Choza Formations, Clear Fork Group, Texas, USA
Specimens:
Length (skull):
Length (total): 328 cm
Weight: around 166 kg
Diet: other tetrapods
Comments: A large representative of the short-skulled lineage of dimetrodonts.  Possibly ancestral to D. angelensis. Although similar in size to Dimetrodon grandis, it differs in the shape of the neural spines, the length of the vertebrae centra, and the lighter overall build. D. gigashomogenes would seem to be a descendent, or possibly even the same species, of the preceding D. dollovanus, from which it differs mainly in larger size.
References: Romer and Price 1940, Reisz 1986




page by M. Alan Kazlev 2002
page uploaded 20 May 2002
checked ATW050514
(originally uploaded on Kheper Site 28 May 1998)
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