Palaeos		Odonatoptera
Arthropoda		Dendrogram

Odonatoptera Dendrogram

PTERYGOTA
|--Palaeodictyopteroida  
|--Ephemeroptera
|--NEOPTERA
|  
`--ODONATOPTERA
   |--Eugeropteridae
   `--Holodonata
      |--Meganisoptera
      `--Odonata

The following dendrogram is combines the material on the old Palaeos edition with a newer but only partial dendrogram by Christopher Taylor. . MAK120516

Odonatoptera Dendrogram

ODONATOPTERA [Odonata sensu lato]  [= Libellulida, Libellulidea, Meganeurina, Odonatoidea, Odonatomorpha]
|  i. s.: Hemeroscopus [Hemeroscopidae] R02
|           `--H. baissicus Pritykina 1977 R02
|         Eocordulia cretacea R02
|         Erasipteroides [Erasipteridae] GE05
|           `--E. valentini GE05
|         Namurotypus [Namurotypidae] RP02
|         Titanophasma fayoli RP02
|         Parahemiphlebia mickoleiti GE05
|--o Eugeropteridae [GEROPTERA] GE05
|    |--Eogeropteron lunatum RP02
|    `--Geropteron arcuatum RP02
`--o NEODONATOPTERA
   |--o "EOMEGANISOPTERA"
   |  `--"Erasipteridae"
   `--o HOLODONATA GE05 (= Euodonatoptera)
      |--o MEGANISOPTERA [Protodonata, Meganeuroidea, Meganeuromorpha] GE05
      |  |--Namurotypidae
      |  `--o MEGANEUROMORPHA (Meganeura)
      |    |--Paralogidae GE05
      |    |--Kohlwaldiidae NB01
      |    |--Kargalotypus Rohdendorf 1962 [Kargalotypidae, Kargalotypinae] NB01
      |    |    `--*K. kargalensis (Martynov 1932) [=Megatypus kargalensis] NB01
      |    `--Meganeuridae GE05
      |         |--Arctotypus sinuatus GE05
      |         |--Meganeuropsis permiana Z02 [incl. M. americana GE05]
      |         |--Meganeura monyi GE05
      |         |--Megatypus schucherti GE05
      |         `--Tupus gracilis GE05
      `--o ODONATOCLADA RP02
         |--o Campylopteridae [Campylopterodea] RP02
         `--Panodialata NB01
            |--Lapeyria [Lapeyriidae] NB01
            |    `--L. magnifica Nel, Gand & Garric 1999 NB01
            `--o NODIALATA RP02
               |--o PROTANISOPTERA  (= Pandiscoidalia RP02)  (alternative (and less resolved) phylogeny)
               |  |--Polytaxineuridae
               |  `--o DITAXINEUROMORPHA
               |     |--Permaeschnidae
               |     `--o DITAXINEURIDA
               |        |--Callimokaltaniidae
               |        `--o DITAXINEUROIDEA
               |           |--Hemizygopteridae
               |           `--Ditaxineuridae
               `--o DISCOIDALIA
                  |--o TRIADOPHLEBIOPTERA
                  |  |--o TRIADOTYPOMORPHA
                  |  |  |--Triadotypidae
                  |  |  `--Piroutetiidae
                  |  `--o TRIADOPHLEBIOMORPHA
                  |     |--o ZYGOPHLEBIOIDEA
                  |     |  |--Xamenophlebiidae
                  |     |  `--Zygophlebiidae
                  |     `--o TRIADOPHLEBIIDA
                  |        |--Mitophlebiidae
                  |        `--o TRIADOPHLEBIOIDEA
                  |           |--Triadophlebiidae
                  |           `--Paurophlebiidae
                  `--o STIGMOPTERA
                     |--o PROTOZYGOPTERA
                     |  |--o PERMAGRIONOIDEA
                     |  |  |--Permagrionidae
                     |  |  `--Permolestidae
                     |  `--o ARCHIZYGOPTERA
                     |     |--Permepallagidae (sedis mutabilis)
                     |     |--Kennedyidae (sedis mutabilis)
                     |     `--o PROTOMYRMELEONTOIDEA (sedis mutabilis)
                     |        |--Batkeniidae
                     |        `--Protomyrmeleontidae
                     `--o PANODONATA
                        |--Tarsophlebiidae
                        `--o ODONATA sensu stricto (modern dragonflies)
                           |-- EPIPROCTOPHORA
                           `-- ZYGOPTERA

[1] Alternative classification:

                        `--Ditaxineuroidea [Protanisoptera] RP02
                             |  i. s.: Hemizygopteron Zallesskiy 1955 NB01
                             |--Ditaxineuridae GE05
                             |--Permaeschna Martynov 1931 NB01 [Permaeschnidae GE05]
                             |--Polytaxineura Tillyard 1935 NB01 [Polytaxineuridae RP02]
                             |--Callimokaltaniidae RP02
                             `--Kaltanoneuridae RP02

*Type species of generic name indicated

Reference(s):

Bechly, G., 1998: Phylogenetic classification of fossil and extant odonates. --iNet: Phylogenetic Systematics of Odonata

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 6: Selkärangattomat.

[NB01] Nel, A., O. Bethoux, G. Bechly, X. Martínez-Delclòs & F. Papier. 2001. The Permo-Triassic Odonatoptera of the "protodonate" grade (Insecta: Odonatoptera). Annales de la Société Entomologique de France (N.S.) 37 (4): 501-525.

[R02] Rasnitsyn, A. P. 2002. Special features of the study of fossil insects. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 8-12. Kluwer Academic Publishers: Dordrecht.

[RP02] Rasnitsyn, A. P., & L. N. Pritykina. 2002. Superorder Libellulidea Laicharting, 1781. Order Odonata Fabricius, 1792. The dragonflies. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 97-104. Kluwer Academic Publishers: Dordrecht.

Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980. Alkuperäisteos: Beaut‚ Du Monde Animal XI & XII Invertébrés. Rizzoli Editore, Milano 1968.

[Z02] Zherikhin, V. V. 2002. Ecological history of the terrestrial insects. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 331-388. Kluwer Academic Publishers: Dordrecht.