Cisuralian Epoch
Paleozoic:
Permian Period
Artinskian Age

The Artinskian Age

The Artinskian Age of the Cisuralian Epoch: 284 to 276 million years ago

Paleozoic Era
   Cambrian Period
   Ordovician Period
   Silurian Period
   Devonian Period
   Carboniferous Period
   Permian Period
      Cisuralian Epoch
         Asselian Age
         Sakmarian Age
         Artinskian Age 
         Kungurian Age
      Guadalupian Epoch
      Lopingian Epoch 

Introduction
Bestiary
Tetrapod Faunas
Tetrapods
   Arroyo Formation

Introduction

Permian scene

As the climate became drier, the early Permian semiaquatic and lowland floodplain Dimetrodon and Eryops dominated fauna shown above, which had flourished for some 25 million years, was replaced by the more advanced floodplain and upland caesid and therapsid fauna.

Known Occurrence of early Permian Tetrapod Faunas

Early Permian tetrapod provinces

This map shows the three known tetrapod provinces (shaded) of the early Permian - the tropical Edaphosaurid (centre), the Kazakhstan Seymouriamorph province (upper right), and the high latitude Gondwana Mesosaurid province (bottom)

From A.R. Milner, "Biogeography of Palaeozoic Tetrapods" fig.13.3; in J.A. Long (ed.) Palaeozoic Vertebrate Biostratigraphy and Biogeography, 1993, John Hopkins University Press, Baltimore

Age

Tetrapod zone

Time*

Location

USA

Western Europe

Eastern Europe

Brazil

southern Africa

India

Texas

Oklahoma

Germany

Russian platform

Parana Basin

South Africa / South-West Africa

Himalayas

Biotic province

Edaphosaurid province 
(equatorial Euramerica)

 

 

Mesosaurid province
(south-west Gondwanaland)

 

Baigendzhinian

10

277

 

278

 

279

Choza

Hennesey

 

 

 

 

 

9

Vale

 

 

 

 

 

8

Arroyo

Garber

 

zone 0

 

 

 

Aktasinian

7

280

 

281

 

282

 

283

Ludens

West Grandfield

 

 

 

 

 

6

Clyde

Deep Red Run

 

 

Irati

White band

 

5

Belle Plains

Wellington formation

 

 

 

 

 

This table is based on J. M. Anderson & A. R. I. Cruikshank, "The Biostratigraphy of the Permian and Triassic, Part 5, a review of the classification and distribution of Permo-Triassic Tetrapods", in Paleontologica Africana, 21, 15-44 (1978); slightly modified.


* approximate time in MYA (millions of years ago) - nearest million year intervals

An Artinskian bestiary

Early Permian Bestiary - Edaphosaurid-Nectridean Province

Illustration by Robert Bakker

Some Non-Tetrapods

Xenacanthus

Xenacanthus


Class Chondrichthyes
Subclass Elasmobranchii
Order Xenacanthida
Family Xenacanthidae

Standard total length of about 1 to 2 meters

Predator of aquatic vertebrates


Some Tetrapods

Aktasinian (Early Artinskian)

Eryops megacephalus [Cope]


Class Tetrapoda
Order Temnospondyli
Superfamily Eryopoidea
Family Eryopidae

Wichita Series (Wolfcampian age), Texas Red Beds

Standard total length from 1.5 to 2 meters

about 130 kg

Predator of other tetrapods

a large long-lived aquatic temnospondyl


Arroyo Formation - Early Baigendzhinian Age

Dimetrodon grandis (Case, 1907)


Plesion Synapsida (Theropsida)
Order Pelycosauria
Suborder Eupelycosauria
Family Sphenacodontidae

Arroyo Formation, Clear Fork Group

Texas, USA

Skull measures 42 cm long. Standard total length of 3.2 metres

around 250 kg

Predator of other tetrapods

The last and largest of the long-skulled, stocky-bodied lineage of dimetrodonts. Apart from D. angelensis this was the largest species of Dimetrodon. A large, heavily-built, very common species, it is close to the earlier D. limbatus, with which it differs in larger size, more elongate and stouter neural spines, and fewer premaxillary teeth (only two on either side, the fewest for any species of the genus). It is associated with the slightly smaller but equally successful Dimetrodon gigashomogenes

Romer and Price 1940, Reisz 1986


Dimetrodon gigashomogenes Case, 1907


Plesion Synapsida (Theropsida)
Order Pelycosauria
Suborder Eupelycosauria
Family Sphenacodontidae

Arroyo, Vale and Choza Formations, Clear Fork Group

Texas, USA

Standard total length of 328 cm

around 166 kg

Predator of other tetrapods

A large representative of the short-skulled lineage of dimetrodonts. Possibly ancestral to D. angelensis. Although similar in size to Dimetrodon grandis, it differs in the shape of the neural spines, the length of the vertebrae centra, and the lighter overall build. D. gigashomogenes would seem to be a descendent, or possibly even the same species, of the preceding D. dollovanus, from which it differs mainly in larger size.

Romer and Price 1940, Reisz 1986


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page by M. Alan Kazlev 2002
page uploaded 20 May 2002
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(originally uploaded on Kheper Site 28 May 1998)
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